A história se desenrola em uma atmosfera sombria do fim dos anos 80, em uma pequena cidade claramente dividida entre liberais e conservadores. Nesse turbilhão se encontra Donnie Darko (Jake Gyllenhaal), um garoto considerado problemático com alguns traços de esquizofrenia (assim caracterizado pela psiquiatra que ele frenquenta, Ms. Thurman). Em uma noite, um coelho monstro gigante acorda Donnie, salvando sua vida, pois repentinamente uma turbina de avião despenca do céu caindo exatamente na cama de Donnie. O coelho monstro gigante ainda profetiza que o mundo irá se acabar dentro de pouco tempo, este mundo, Donnie entenderá ser o mundo pessoal dele. Donnie se mostra dividido entre a realidade e suas alucinações, junto a isso, muitos questionamentos sobre o sentido da vida e, principalmente, da morte.
O filme aborda temas da física teórica como a possibilidade de viagem ao tempo usando obras como de Stephen Hawking - Uma Breve História do Tempo e Roberta Sparrow - Filosofia da Viagem do Tempo para expor tais ideias.
O filme trabalha a capacidade da pessoa trabalhar o que é o mundo de verdade, o lugar em que ele se encontra, ou o pessoal da própria vida.
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